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Below are the restricted articles (PDF files) that you requested, for your personal use only. They must not be copied or sold, and this link must not be transmitted to anyone.

  1. Luedtke J, et al. 2023. Ongoing declines for the world's amphibians in the face of emerging threats. Nature 622: 308-314. E-print
  2. Craig J, Kumar S, Hedges SB. 2023. The origin of eukaryotes and rise in complexity were synchronous with the rise in oxygen. Frontiers in Bioinformatics DOI: 10.3389/fbinf.2023.1233281. E-print
  3. Hedges SB. 2023. Book Review: the conservation and biogeography of amphibians in the Caribbean. Phyllomedusa 22: 87-93. E-print
  4. Ali J, Hedges SB. 2023. The colonisation of Madagascar by land-bound vertebrates. Biological Reviews 98(5): 1583-1606. E-print
  5. Hedges SB. 2022. A new species of frog from the Caribbean island of Montserrat(Eleutherodactylidae, Eleutherodactylus). Zootaxa 5219: 375-387. E-print
  6. Landestoy MA, Schools M, Hedges SB. 2022. A new genus and species of Caribbean forest lizard (Diploglossidae; Celestinae)from southern Hispaniola. Zootaxa 5219: 201-226. E-print
  7. Ali J, Hedges SB. 2022. A review of geological evidence bearing on proposed Cenozoic land connections between Madagascar and Africa and its relevance to biogeography. Earth-Science Reviews. Published online 7 July 2022 E-print
  8. Schools M, Kasprowicz A, Hedges SB. 2022. Phylogenomic data resolve the historical biogeography and ecomorphs of Neotropical forest lizards (Squamata, Diploglossidae). Molecular Phylogenetics Evolution 175:107577. E-print
  9. Diaz LM, Fong A, Salas D, Hedges SB. 2021. A new semifossorial snake of the genus Arrhyton (Squamata: Dipsadidae) from eastern Cuba, with taxonomic comments on other species. Zootaxa 5052:406-418. E-print
  10. Moore RD, Hedges SB. 2021. Private conservation of remnant forest ecosystems to support sustainable development: Grand Bois privately protected area, Haiti. Pp. 50-53 in Kettunen M, Dudley N, Gorricho J, Hickey V, Krueger L, MacKinnon K, Oglethorpe J, Paxton M, Robinson JG, Sekhran N, Building on nature: area-based conservation as a key tool for delivering SDGs (IEEP, IUCN WCPA, The Nature Conservancy, The World Bank, UNDP, Wildlife Conservation Society and WWF). E-print
  11. Schools M, Hedges SB. 2021. Phylogenetics, classification, and biogeography of the Neotropical forest lizards (Squamata, Diploglossidae). Zootaxa 4974:201-257. E-print
  12. Chapple DG, Roll U, Bohm M, Aguilar R, Amey AP, Austin CC, Baling M, Barley AJ...Hedges, SB...et al. 2021. Conservation status of the worlds skinks (Scincidae): taxonomic and geographic patterns in extinction risk. Conservation Biologyhttps://doi.org/10.1016/j.biocon.2021.109101. (published online). E-print
  13. McCranie JR, Matthews AJ, Hedges SB. 2020. A morphological and molecular revision of lizards of the genus Marisora (Squamata: Mabuyidae) from Central America and Mexico, with descriptions of four new species. Zootaxa 4763: 301-353. E-print
  14. Hedges SB. 2020. Molecular clocks, flotsam, and Caribbean islands. Pp. 349-351 in Cox, CB, Laddle RJ, & Moore PD, Biogeography: an ecological and evolutionary Approach. 10th Edition (John Wiley & Sons, Inc., Hoboken, New Jersey). E-print
  15. Köhler G, Zimmer C, McGrath K, Hedges SB. 2019. A revision of the genus Audantia of Hispaniola with description of four new species (Reptilia, Squamata, Dactyloidae). Novitates Caribaea. 14: 1-104. doi/10.33800/nc.v0i14.201 E-print (6MB) E-print (55MB)
  16. Hedges SB, Cohen WB, Timyan J, Yang Z. 2018. Reply to Wampler et al.: Deforestation and biodiversity loss should not be sugarcoated. Proc. Natl. Acad Sci. USA. 116: doi/10.1073/pnas.1901879116. E-print
  17. Rapacciuolo G, Graham CH, Marin J, Behm JE, Costa GC, Hedges SB, Helmus MR, Radeloff VC, Young BE, Brooks TM. 2018. Species diversity as a surrogate for conservation of phylogenetic and functional diversity in terrestrial vertebrates across the Americas. Nature Ecol. Evol. 2(12):10.1038/s41559-018-0744-7. E-print
  18. Hedges SB, Cohen WB, Timyan J, Yang Z. 2018. Haiti's biodiversity threatened by nearly complete loss of primary forest. Proc. Natl. Acad Sci. USA. 115(46):11850-11855. E-print Supp
  19. Hedges SB, Tao Q, Walker M, Kumar S. 2018. Accurate timetrees require accurate calibrations. Proc. Natl. Acad Sci. USA. 115(41):E9510-E9511. E-print
  20. Miralles A, Marin J, Markus D, Herrel A, Hedges SB, Vidal N. 2018. Molecular evidence for the paraphyly of Scolecophidia and its evolutionary implications. J Evol Biol. doi: 10.1111/jeb.13373. E-print
  21. Marin J, Hedges SB. 2018. Undersampling genomes has biased time and rate estimates throughout the tree of life. Mol. Biol. Evol. 35: 2077–2084. E-print Corrigendum
  22. Landestoy MA, Turner DB, Marion AB, Hedges SB. 2018. A new species of Caribbean toad (Bufonidae, Peltophryne) from southern Hispaniola. Zootaxa, published online 5 April 2018. doi: 10.11646/zootaxa.4403.3.6. E-print
  23. Marin J, Rapacciuolo G, Costa GC, Graham CH, Brooks TM, Young BE, Radeloff VC, Behm JE, Helmus MR, Hedges SB. 2018. Evolutionary time drives global tetrapod diversity. Roy Soc. Proc B, published online 7 February 2018. doi: 10.1098/rspb.2017.2378. E-print
  24. Holt BG, Costa GC, Penone C, Lessard J-P, Brooks TM, Davidson AD, Hedges SB, Radeloff VC, Rahbek C, Rondinini C, Graham CH. 2017. Environmental variation is a major predictor of global trait turnover in mammals. J. Biogeography 2017: 1-13. E-print
  25. Heinicke MP, Lemmon AR, Lemmon EM, McGrath K, Hedges SB. 2017. Phylogenomic support for evolutionary relationships of New World direct-developing frogs (Anura: Terraranae). Molec. Phylogenet. Evol. 118: 145-155. E-print
  26. Brum FT, Graham CH, Costa GC, Hedges SB, Penone C, Radeloff VC, Rondinini C, Loyola R, Davidson AD. 2017. Global priorities for conservation across multiple dimensions of mammalian diversity. Proc. Natl. Acad. Sci. USA. 114(29): 7641–7646. E-print
  27. Kumar S, Stecher G, Suleski M, Hedges SB. 2017. TimeTree: A resource for timelines, timetrees, and divergence times. Mol. Biol. Evol. doi: 10.1093/molbev/msx116. E-print
  28. Tucker DB, Hedges SB, Colli GR, Pyron RA, Sites JW, Jr. 2017. Genomic timetree and historical biogeography of Caribbean island ameiva lizards (Pholidoscelis: Teiidae). Ecology and Evolution. doi: 10.1002/ece3.3157. E-print
  29. Rapacciuolo G, Marin J, Costa GC, Helmus MR, Behm JE, Brooks TM, Hedges SB, Radeloff VC, Young BE, Graham CH. 2017. The signature of human pressure history on the biogeography of body mass in tetrapods. Global Ecology and Biogeography. doi: 10.1111/geb.12612. E-print Not Available
  30. Marin J, Battistuzzi FU, Brown AC, Hedges SB. 2017. The Timetree of Prokaryotes: New Insights into Their Evolution and Speciation. Mol. Biol. Evol. doi: 10.1093/molbev/msw245. E-print
  31. Penone C, Weinstein BG, Graham CH, Brooks TM, Rondinini C, Hedges SB, Davidson AD, Costa GC. 2016. Global mammal beta diversity shows parallel assemblage structure in similar but isolated environments. Proc. R. Soc. B 283: 20161028. doi: 10.1098/rspb.2016.1028. E-print
  32. Duellman WE, Marion AB, Hedges SB. 2016. Phylogenetics, classification, and biogeography of the treefrogs (Amphibia: Anura: Arboranae). Zootaxa 4104: 1-109. E-print
  33. Marin J, Hedges SB. 2016. Time best explains global variation in species richness of amphibians, birds and mammals. J. Biogeography 43: 1069-1079. E-print Supp1 Supp2
  34. Nagy ZT, Marion AB, Glaw F, Miralles A., Noppers J, Vences M, Hedges SB. 2015. Molecular systematics and undescribed diversity of Madagascan scolecophidian snakes (Squamata: Serpentes). Zootaxa 4040: 031–047. E-print
  35. Schmid M et al. (104 authors) 2015. Third report on chicken genes and chromosomes. Cytogenet Genome Res. 145:78-179 E-print
  36. Heinicke MP, Barrio-Amorós CL, and SB Hedges. 16 Jul 2015. Molecular and morphological data support recognition of a new genus of New World direct-developing frog (Anura: Terrarana) from an under-sampled region of South America​. Zootaxa 3986 (2): 151–172. E-print
  37. Hedges SB. 2014. The high-level classification of skinks (Reptilia, Squamata, Scincomorpha). Zootaxa 3765: 317-338. E-print
  38. Hedges SB. 2014. The origin of life. In Oxford Bibliographies in Evolutionary Biology Ed. Jonathan Losos. New York: Oxford University Press. E-print
  39. McCranie JR, Hedges SB. 2013. A review of the Cnemidophorus lemniscatus group in Central America (Squamata: Teiidae), with comments on other species in the group. Zootaxa 3722: 301-316. E-print
  40. McCranie JR, Hedges SB. 2013. A new species of Phyllodactylus (Reptilia, Squamata, Gekkonoidea, Phyllodactylidae) from Isla de Guanaja in the Honduran Bay Islands. Zootaxa 3694: 051-058. E-print
  41. McCranie JR, Hedges SB. 2013. Two additional new species of Sphaerodactylus (Reptilia, Squamata, Gekkonoidea, Sphaerodactylidae) from the Honduran Bay Islands. Zootaxa 3694: 040-050. E-print
  42. Hedges SB. 2013. Revision shock in taxonomy. Zootaxa. Zootaxa 3681: 297-298. E-print
  43. Hedges SB. 2013. Caribmap: a mobile-friendly web resource for Caribbean maps that facilitates quantitative historical analysis. Journal of Map and Geography Libraries 9(3): 350-360. pre-print
  44. (this is the accepted manuscript; see journal website for tightly-controlled published version).
  45. Barrio-Amoros CL, Heinicke MP, Hedges SB. 2013. A new tuberculated Pristimantis (Anura, Terrarana, Strabomantidae) from the Venezuelan Andes, redescription of Pristimantis pleurostriatus, and variation within Pristimantis vanadisae. Zootaxa. Zootaxa 3647: 43-62. E-print
  46. Hedges SB. 2013. Wormholes record species history in space and time. Biology Letters dx.doi.org/10.1098/rsbl.2012.0926. Published online 21 November 2012, in print 23 February 2013. E-print Supplement
  47. Marin J, Donnellan SC, Hedges SB, Doughty P, Hutchinson MN, Cruaud C, Vidal N. 2013. Tracing the history and biogeography of the Australian blindsnake radiation. J. Biogeogr. 40: 928-937, published online 23 November 2012. E-print
  48. Supp1 Supp2
  49. McCranie JR, Hedges SB. 2012. Two new species of geckos from Honduras and resurrection of Sphaerodactylus continentalis Werner from the synonymy of Sphaerodactylus millepunctatus Hallowell (Reptilia, Squamata, Gekkonoidea, Sphaerodactylidae). Zootaxa 3492: 65-76. E-print
  50. Hedges, SB, Conn CE. 2012. A new skink fauna from Caribbean islands (Squamata, Mabuyidae, Mabuyinae). Zootaxa 3288: 1-244. E-print
  51. Díaz, LM, SB Hedges, M Schmid. 2012. A new cryptic species of the genus Eleutherodactylus (Amphibia: Anura: Eleutherodactylidae) from Cuba. Zootaxa 3220: 44-60. E-print
  52. Adalsteinsson, S.A., Branch, W.R., Trape, S., Vitt, L.J., & Hedges, S.B. 2009. Molecular phylogeny, classification, and biogeography of snakes of the Family Leptotyphlopidae (Reptilia, Squamata). Zootaxa 2244: 1-50. E-print
  53. Heinicke, M.P., Duellman, W. E., Trueb, L., Means, D. B., MacCulloch, R. D., Hedges, S. B. 2009. A new frog family (Anura: Terrarana) from South America and an expanded direct-developing clade revealed by molecular phylogeny. Zootaxa 2211: 1-35. E-print
  54. Hedges, S. B., A. Couloux, and N. Vidal. 2009. Molecular phylogeny, classification, and biogeography of West Indian racer snakes of the Tribe Alsophiini (Squamata, Dipsadidae, Xenodontinae). Zootaxa 2067:1-28. E-print
  55. Dίaz, L. M. and S. B. Hedges. 2009. First record of the genus Aristelliger (Squamata: Sphaerodactylidae) in Cuba, with description of a new species. Zootaxa 2028:31-40. E-print
  56. Vidal, N., W.R. Branch, O.S.G. Pauwels, S.B. Hedges, D.G. Broadley, M. Wink, C. Cruaud, U. Joger & Z.T. Nagy. 2008. Dissecting the major African snake radiation: a molecular phylogeny of the Lamprophiidae Fitzinger (Serpentes, Caenophidia). Zootaxa 1945:51–66. E-print
  57. Hedges, S. B., W. E. Duellman, and M. P. Heinicke. 2008. A replacement name for Isodactylus. Hedges, Duellman, and Heinicke, 2008. Zootaxa 1795:67-68. E-print
  58. Dίaz, L. M. and S. B. Hedges. 2008. A new gecko of the genus Tarentola (Sauria: Gekkonidae) from eastern Cuba. Zootaxa 1743:43–52. E-print
  59. Hedges, S. B., W. E. Duellman, and M. P. Heinicke. 2008. New World direct-developing frogs (Anura: Terrarana): molecular phylogeny, classification, biogeography, and conservation. Zootaxa 1737:1–182. E-print
  60. Duellman, W. E. and S. B. Hedges. 2008. Two new minute species of Phrynopus (Lissamphibia: Anura) from the Cordillera Oriental in Peru. Zootaxa 1675:59–66. E-print
  61. Lehr, E., J. B. Pramuk, S. B. Hedges, and J. H. Córdova. 2007. A new species of arboreal Rhinella (Anura: Bufonidae) from Yanachaga-Chemillén National Park in central Peru. Zootaxa 1662:1–14. E-print
  62. Thomas, R. and S. B. Hedges. 2007. Eleven new species of snakes of the genus Typhlops (Serpentes: Typhlopidae) from Hispaniola and Cuba. Zootaxa 1400:1-26. E-print

Reprints of other papers from Hedges Lab are available on this public web site: