Courses.

Undergraduate (every fall semester):

BIOLOGY 427 ­ Evolution: a comprehensive course in evolutionary biology, covering concepts (history, Darwin, etc.), Earth history, the origin of life, fossil record, macroevolution, adaptation, systematics, genes and populations, molecular evolution, molecular phylogenetics, speciation, biogeography, coevolution, primate evolution, biodiversity, and astrobiology. (3 credits)

Graduate (usually each fall semester):

BIOLOGY 514 ­ Topics in Evolution: exploration of recent literature in evolutionary biology. (2 credits)

• Descriptions of graduate courses at Penn State
• Current semester schedule listings of courses

In 2010 the U. S. National Research Council ranked Penn State's research-doctoral program in BIOLOGY 8th in the U.S. among 212 ranked universities.